Because the amount of oil you are losing is excessive, even for an old engine, and from the results of your other tests, I'm beginning suspect one of two things (or a likely combination of both):
1. Little care or attention was given to the assembly of the half moon seal. This is a semi-circular cast iron piece at the back of the block and mates with the rear main cap and completes the function of the scroll seal.
2. Improper assembly of the oil pan seals.
If your engine was assembled with cork seals at the front and rear of the pan (around the front and rear main caps). Many amateur/inexperienced engine builders don't realize the cork gaskets lengths are "run wild" and either leave them too long, which leaves a gap for oil to escape; or the were trimmed flush, preventing a good seal. The end result is the same. Some gasket sets include formed neoprene seals which do not have the same problem. Unfortunately, there is no easy way to see what's in there short of dropping the pan. Another issue is if the pan bolt were over torqued, which distorts the flange on the pan.
If all else fails, I would drop the oil pan to examine the seals, but if it's the half moon seal that is the culprit, there's no choice but to pull the engine.